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LITERATURE CITED

Bolen, E. G. and W. L. Robinson. Wildlife Ecology and Management. Prentice Hall, Inc. Englewood Cliffs, New Jersey. 620 pp. Lftner, David C., J.A. Shuey, and J.V. Calhoun. 1992. Butterflies and Skippers of Ohio. Ohio Biol. Surv. Bull. New Series Vol. 9 No. 1 xii + 212 p. (includes 40 Pls.). Missouri Department of Conservation. 1990. Butterfly Gardening and Conservation. 1990. Urban Wildlife Series, Pamphlet no. 2. NH-4/90-15M. Mooberxy, F.M. 1996. The Billiance of Asters: Part 1. The East. American Butterflies. Vol. 4(3): 22-27. Opler, Paul A. 1992. Eastern Butterflies. Peterson Field Guides, Houghton Muffin Company, New York. 396 pp. Pullin, Andrew S., Ed. 1995. Ecology and Conservation of Butterflies. The British Butterfly Conservation Society. Chapman and Hall, London. 363 pp. Primack, Richard B. A Primer of Conservation Biology. 1995. Sinauer Associates, Inc., Sunderland, Mass. 277 pp. Shull, Ernest M. Tekulsky, Mathew. 1987. The Butterflies of Indiana. Indiana Academy of Science. 1985. The Butterfly Garden. 262 pp. The Harvard Common Press, Boston, Massachussets.144 pp. The Xerces Society. Institution. 1990. Butterfly Gardening.
Sierra Club Books, San Francisco. Created in association with The Smithsonian 192 pp.



 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 




NETTLE LAKE BUTTERFLY RESTORATION/CONSERVATION PLAN


     Most systematists recognize the taxonomic label “butterflies” to be composed of two Hesperioidea..........Click here for the larger image.... Super families of insects, the “true butterflies",” the Papilionoidea", and the “skippers,” the Hesperioidea. Skippers tend to be small to medium-sized, drab in color, with brown, orange, and black as frequent colors, and have hooked antennal clubs, in contrast to “true” butterflies. Their flight is often erratic, and they have been described as resembling miniature fighter planes. Unless indicated, henceforth the term “butterfly” will refer to an individual of either super family. Like 88% of the insects, butterflies undergo a complete metamorphosis, with 4 stages to their life cycle - egg, larva (caterpillar), pupa (chrysalis), and adult. Most species also have a resting stage, diapause, so caterpillars hatch having something to eat, and adults emerge at a favorable time. In most cases, larval food plants differ from adult nectar plants, which is thought to reduce competition between caterpillars and adults. Thus, to support minimum viable populations of a particular species, having plants suitable for both life cycle stages is a necessity.
     This restoration plan is aimed at maximizing butterfly diversity at Nettle Lake, and additionally, targeting species designated endangered, threatened, or of special interest by the Ohio Department of Natural Resources Division of Wildlife. Currently four species that have been documented in Williams County are so designated: Two-Spotted Skipper, Karner Blue Purplish Copper, Regal Fritillary, and Silver-Boardered Fritillary. Additionally, the Karner Blue butterfly (pictured to the right), Lycae ides melissa samuelis, currently designated as Federally Endangered, has been documented in the Oak Openings of Lucas County, and in northern Indiana as far east as Lake County, although it has not been documented in Williams County.
      Butterflies are probably some of our most recognizable and charismatic invertebrates, and currently much restoration work is aimed at preserving them because of their aesthetic values (i.e., the current rising popularity of “butterfly gardening”) Additionally, however, butterflies may serve a more valuable function, as biological indicators of ecosystem health and function. Click here for more information on Butterflies.

    Of the 524 species of butterflies and skippers found in the eastern United States, 144 have been documented in Ohio. A list of species possibly occurring at Nettle Lake was compiled by examining published accounts of species distributions in both Indiana and Ohio. A species was included in the list if it has been collected in either Williams County, Ohio, or Steuben County, Indiana. Eighty-three species comprised this list. Data on existing species at Nettle Lake were compiled by two ways: sightings recorded by volunteer landowners, and specimens collected or sighted by student field assistants, from May 9, 1995, to Oct. 31, 1996. Only thirteen species were documented as currently known to occur at this site, leaving 70 species of possible occurrence at Nettle Lake. The relatively low number of species observed is probably both a result of observer inexperience and the relatively low number of observer-days censusing butterflies only (i.e., in some cases no record in the Biological Survey probably does not reflect that the species is truly absent). The Biological Survey also documented tree and wildflower species present. Records of known larval host plants and adult nectar sources were compared to existing vegetation at the site to identify any “gaps” in habitat requirements for butterflies potentially using the area around Nettle Lake. Additionally, attention was given to identifying any special habitat requirements of the four state endangered or threatened species on the list.

    The host plants of many of the skippers are unknown in Ohio. Twenty of the 29 species (68%) of skippers possibly occurring at Nettle Lake use several species of grasses and sedges in other parts of their range. Species commonly used are: purple-top, little bluestem, meadow grasses, and panic grasses. Of these four species, Andropogon is the only one not relatively common throughout the state; thus habitat enhancement for skippers at this site might include developing a stand of this species for larval food needs. However, additional information on which species of grasses and sedges skippers are known to use in Ohio would be helpful, in any future monitoring program.

    Most adult skippers use a diversity of nectar plants. Nettle Lake has a variety of the plants commonly used: red clover, alfalfa, Canada thistle, raspberry, swamp milkweed, teasel, wild bergamot, spring beauty, common dandelion, and oxeye daisy. However, some of the recommendations below may enhance nectaring habitat quality and quantity for adult skippers.

    Very broadly speaking, the true butterflies tend to be more specific in their required host plants than their nectar plants. Common nectar plants used are similar to the ones used by skippers. Some groups that deserve mention, because of their particularly specialized needs, or because Nettle Lake is either scarce in or lacking their host plants: the Swallowtails, the Fritillaries, other Nymphalids, and the Satyrs and the Wood Nymphs.

   Mourning CloakSome forest butterfly species visit patchy resources such as rotting fruit, fermenting sap flows, dung, or carrion for their primary adult energy resources, and may infrequently visit flowers. Consequently, these species may be hard to manage for adult needs on a consistent basis. This list includes the Question Mark, Comma, Gray Comma, Compton Tortoise Shell, and Mourning Cloak.




Butterfly and skipper species documented in Williams Co. Ohio, and Steuben Co., Indiana, and occurring or potentially occurring at Nettle Lake. (red type print indicates those species observed in the Biological Survey).

SUPERFAMILY Hesperioidea Latreille - Skippers

  • FAMILY Hesperiidae - True Skippers
    Silver-spotted Skipper
    Southern Cloudy Wing
    Northern Cloudy Wing
    Sleepy Dusky Wing
    Juvenal’s Dusky Wing
    Columbine Dusky Wing
    Checkered Skipper
    Common Sooty Wing
    Least Skipper
    European Skipper
    Fiery Skipper
    Leonard’s Skipper
    Peck’s Skipper
    Tawny-Edged Skipper
    Cross Line Skipper
    Long Dash
    Northern Broken Dash
    Little Glassy Wing
    Sachem
    Delaware Skipper
    Mulberry Wing
    Northern Golden Skipper
    Broad-winged Skipper
    Dion Skipper
    Duke’s Skipper
    Black Dash
    Two-spotted SkipperE
    Dun Skipper
    Ocala Skipper


    SUPERFAMILY Papilionoidea Latreille - True Butterflies


  • FAMILY Papilionidae - Swallowtails
    Pipevine Swallowtail
    Zebra Swallowtail
    Black Swallowtail
    Giant Swallowtail
    Tiger Swallowtail
    Spicebush Swallowtail

  • FAMILY Pieridae - Whites and Sulfurs
    Checkered White
    European Cabbage White
    Clouded Sulfur/Common Sulfur Alfalfa Butterfly/Orange Sulfur
    Dog Face
    Little Sulfur
    Sleepy Orange

  • FAMILY Lycaenidae - Gossamer Wings
    American Copper
    Bronze Copper
    Purplish CopperE
    Dorcas copper
    Coral Hairstreak
    Acadian Hairstreak
    Banded Hairstreak
    Hickory Hairstreak
    Striped Hairstreak
    Eastern Tailed Blue
    Spring Azure

  • FAMILY Libytheidae - Snouts 
    Eastern Snout Butterfly

  • Great SpangledFAMILY Nymphalidae - Brushfoots
    Variegated Fritillary
    Great Spangled Fritillary
    Aphrodite Fritillary
    Regal FritillaryE
    Silver-bordered FritillaryT
    Meadow Fritillaiy
    Silvery Checkerspot
    Pearl Crescent
    Baltimore
    Question Mark
    Mourning Cloak Comma
    Gray Comma
    Compton Tortoise Shell
    Mourning Cloak
    Milbert’s Tortoise Shell
    American Painted Lady
    Painted Lady
    Red Admiral
    White Admiral/Red-Spotted Purple
    Buckeye
    Viceroy

  • FAMILY Apaturidae - Leaf Wing and Hackberry Butterflies
    Hackberry Butterfly
    Tawny Emperor

  • FAMILY Satyridae - Satyrs and Wood Nymphs
    Northern Pearly Eye
    Northern Eyed Brown
    Appalachian Eyed Brown
    Little Wood Satyr
    Common Wood Nymph


  • FAMILY Danaidae - Milkweed Butterflies
    Monarch

    Native Plants 

    Host Plants
    dogbanes
    milkweeds - butterfly weed, 
    purple milkweed, and 
    swamp milkweed.
    willows
    sassafras
    wild carrot and relatives
    violets
    Cultivated Plants

    Host Plants
    carrot, dill, parsley
    asters
    cabbage, broccoli





    Nectar Plants

    milkweeds
    asters
    ox-eye daisy
    coreopsis
    coneflowers
    bergamots
    phlox
    blazing stars
    common lilac

    Nectar Plants

    asters
    butterfly bush
    daisies or chrysanthemums
    hibiscus
    french lavender
    alfalfa
    bee balm
    phlox
    marigolds
    Vinca
    zinnias

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